The site recorded a month-on-month rise of 4.14 per cent to 27,324,209 uniques; while last month's leader Telegraph.co.uk posted a 13.83 per cent drop in unique users to 23,875,251.
The Guardian, Mirror Group and Independent were the only titles to increase unique user numbers in April. Mail Online, Times Online and The Sun all recorded slight drops.
Having taken the top spot in February's ABCes, Sun Online's figures now place it behind News International stablemate Times Online in terms of uniques.
The full figures for the six audited titles and Mirror Group's websites are listed below. The percentage in brackets indicates the month-on-month change compared with March's ABCe report:
Telegraph.co.uk
Unique users: 23,875,251 (-13.83 per cent); +28.04 per cent year-on-year
Guardian.co.uk
Unique users: 27,324,309 (+4.14 per cent); +47.33 per cent year-on-year
Mail Online
Unique users: 23,153,003 (-2.59 per cent); + 28.34 per cent year-on-year
Times Online
Unique users: 21,585,569 (-0.13 per cent); +40.11 per cent year-on-year
TheSun.co.uk
Unique users: 20,567,178 (-3.21 per cent); +46.19 per cent year-on-year
Independent.co.uk
Unique users: 10,432,660 (+3.97 per cent); +63.19 per cent year-on-year
Mirror Group
Unique users: 8,596,963 (+20.44 per cent); +100.98 per cent year-on-year
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